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life-history evolution life historic characteristics life cycle and history reproductive histories life-history strategy life-histories life-history strategists Evolutionary life-history theory life history evolution life history characteristics life-history theories life history strategy life-history studies life history strategies Life History Life_history_theory#Determinants life-history traits life history traits life-history theory life cycle life history theory life history Life history Life History Theory life-history r/K life history theory fitness trade-offs life histories life-history pattern