clgo:title | - Brackenbury Professor of Physiology(1873)
- FRCP Lond. (1896)
- FRCPEdin. (1872)
- FRS(1872)
- Fullerian Professor of Physiology(1884–1886)
- MDEdin.(1862)
- MRCPEdin.(1871)
- MRCPLond.(1885)
- President of the biological section of theBritish Association(1882)
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